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==Systematics== About 40 species have been described from different animals, but many of them are probably [[Synonym (taxonomy)|synonyms]].<ref name="Meyer1979">{{cite journal |author1=Meyer E.A. |author2=Radulescu S. | year = 1979 | title = ''Giardia'' and Giardiasis | journal = Advances in Parasitology | volume = 17 | pages = 1β47 | pmid = 395833 | doi = 10.1016/S0065-308X(08)60548-5 | quote = no |isbn=9780120317172 }}</ref> Currently, five to six morphologically distinct species are recognised.<ref>{{cite book |first1=R.C. |last1=Brusca |first2=G.J. |last2=Brusca |title=Invertebrates |publisher=Sinauer Associates |edition=2nd |year=2003 |isbn=0878930973 }}</ref> ''[[Giardia lamblia]]'' (=''G. intestinalis'', =''G. duodenalis'') infect humans and other mammals, ''G. muris'' is found from other mammals, ''G. ardeae'' and ''G. psittaci'' from birds, ''G. agilis'' from amphibians and ''G. microti'' from [[vole]]s.<ref name=Adam01/> Other described (but not certainly valid), species include:<ref>{{cite web |title=Giardia Kunstler |date=September 2008 |work=Tree of Life Web Project |url=http://tolweb.org/Giardia/97370/2008.09.02}}</ref> {{div col|colwidth=22em}} *''Giardia beckeri'' *''Giardia beltrani'' *''Giardia botauri'' *''Giardia bovis'' *''Giardia bradypi'' *''Giardia canis'' *''Giardia caprae'' *''Giardia cati'' *''Giardia caviae'' *''Giardia chinchillae'' *''Giardia dasi'' *''Giardia equii'' *''Giardia floridae'' *''Giardia hegneri'' *''Giardia herodiadis'' *''Giardia hyderabadensis'' *''Giardia irarae'' *''Giardia marginalis'' *''Giardia melospizae'' *''Giardia nycticori'' *''Giardia ondatrae'' *''Giardia otomyis'' *''Giardia pitymysi'' *''Giardia pseudoardeae'' *''Giardia recurvirostrae'' *''Giardia sanguinis'' *''Giardia serpentis'' *''Giardia simoni'' *''Giardia sturnellae'' *''Giardia suricatae'' *''Giardia tucani'' *''Giardia varani '' *''Giardia viscaciae'' *''Giardia wenyoni'' {{div col end}} Many different species of ''Giardia'' exist, so to differentiate between species, very specific PCR (Polymerase Chain Reactions) have been developed to detect specific ''Giardia'' spp. Gene probe-based detection is also used to differentiate between species of ''Giardia''. A more common and less time-consuming means of identifying different species of ''Giardia'' includes microscopy and immunofluorescence techniques.<ref>{{harvnb|Mahbubani|1992}}</ref> Genetic and biochemical studies have revealed the heterogeneity of ''[[Giardia lamblia]]'', which contains probably at least eight lineages or [[cryptic species]].<ref>{{cite journal |vauthors=Thompson RC, Monis PT |title=Variation in Giardia: implications for taxonomy and epidemiology |journal=[[Adv. Parasitol.]] |volume=58 |pages=69β137 |year=2004 |pmid=15603762 |doi=10.1016/S0065-308X(04)58002-8 |series=Advances in Parasitology |isbn=9780120317585 }}</ref>
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